Chemically, cellulose is a linear polymer that is composed of D-glucose subunits linked by β-1, 4 glycosidic bonds forming the dimer cellobiose. These form long chains (or elemental fibrils) linked together by hydrogen bonds and van der Waals forces. Cellulose usually is present as a crystalline form and a small amount of no organized cellulose chains forms amorphous cellulose. In the latter conformation, cellulose is more susceptible to enzymatic degradation ( Perez et al ., 2002 ). Cellulose appears in nature to be associated with other plant compounds and this association may affect its biodegradation.